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Years ago, Guy’s parents get trapped in tar. As they sink into it, they tell him to find somewhere called ‘tomorrow’. He goes on a long journey and meets a young Belt before eventually coming across Eep, leading to the events of the first film.
The Croods, along with Guy and their pets Chunky and Douglas, are still searching for a place to settle down, all the while Grug acts annoyed at Eep and Guy’s blossoming romance. One night, Grug comes across a giant wall and leads the whole pack to it. They are soon caught in a net and are released by the owners of the land, a couple named Phil and Hope Betterman and they are soon revealed to be friends of Guy’s late parents and therefore, know Guy. The Bettermans welcome the Croods to their giant tree-home as house guests, where they meet their daughter and Guy’s old friend Dawn who immediately befriends Eep. Life with the Bettermans becomes degrading for Grug due to the Bettermans being better evolved. Simultaneously, the Bettermans are revealed to have prejudice towards cave people and believe that Guy is better off with them because of his greater evolution, so they hatch a scheme to get Guy to hook up with Dawn and to leave the Croods. Phil eventually takes Grug to his secret man-cave, a sauna-like place behind the waterfalls, where he manipulates him into believing Guy should leave their pack, and Hope gets on Ugga’s bad side by demeaning her family’s nature as cave people. During this, Guy immediately becomes accustomed to life with the Bettermans, adjusting to their ways but unintentionally drifting from Eep in the process.
Meanwhile, Eep discovers that Dawn has never been outside the wall, and, connecting this to the solitude she faced in her cave, uses Chunky to escape the land with Dawn and jump their wall for a joyride that ends with a bee stinging Dawn and having her hand swell up. When Eep takes her back home, Guy, upon finding out, chides her for her recklessness, ending with him insensitively calling her a ‘cave-girl’. At dinner, tensions rise between the parents, Guy and Eep, especially when Dawn’s swelling is revealed. Having enough, the Croods decide to leave, and Guy and Eep have a huge falling out, which results in Guy’s decision to stay with the Bettermans. Grug reveals that he and Ugga have eaten an entire supply of bananas hoarded around the Bettermans land that Phil had forbidden him from consuming. This causes the land to be attacked by ‘Punch-Monkeys’, monkeys with human-like strength, whom Phil had sent the bananas to every day so that they would leave his family alone. Upset over the lack of bananas, the Punch-Monkeys kidnap Grug, Phil, and Guy and take them to their homeland.
As the men are taken, the women, Thunk, and Chunky attempt to rescue them. During their time together, Hope vents out her frustrations with the Croods but grows to accept the Croods after she gets marooned on an island full of Wolf-Spiders, and the family makes peace with them. They eventually form a team called the Thundersisters, which was an old girl-group Gran was in when she was younger, to mount a rescue. At the Punch-Monkey home, Grug, Guy, and Phil soon discover that Phil unintentionally deprived the Punch-Monkeys of their water resource and that the Punch-Monkeys need the bananas not only to eat themselves, but to offer to a tyranical Spiny Mandrilla in hopes of appeasing its appetite. The monkeys make Grug and Phil fight gladiator-style to see who will be the sacrifice and when they wear each other out, they exchange their bitter feelings with each other, making Guy realize and regret the effects his words had on Eep.
Soon, the monkeys dress all three men as bananas to sacrifice to the giant Spiny Mandrilla. Grug and Phil apologize for their poor behavior and for putting pressure on Guy, but just as they are about to be eaten, the Thundersisters show up to rescue them. A long and perilous battle soon ends with Guy and Eep on a giant skull-chandelier where they reconcile and soon use it to defeat the Spiny Mandrilla by using fire to cut the ropes and send it falling into the abyss below when it pulls off Eep’s “peanut toe”, which she used as a prosthetic limb, allowing the families to escape. Afterwards, Eep and Guy share their first kiss.
With their differences finally settled, the Bettermans allow the Croods to live in their land as neighbors along with the monkeys grateful to the humans for killing the Mandrilla, and tear down the wall surrounding their land. Guy declares that Eep is his ‘tomorrow’ and he and Eep soon move into one of the Betterman’s bedrooms together, with Grug finally accepting that his daughter wants to be closer to Guy. The families continues to grow closer to each other, agreeing to Grug’s earlier motto to keep the ‘pack’ together.
streaming online The Croods: A New Age (2020)